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A better understanding and a more quantitative design of mixed-species stands will contribute to more integrative and goal-oriented research in mixed-species forests. Much recent work has indicated that the structure and growth of mixed species forests may fundamentally differ from monocultures. Here we suggest how to progress from the present accumulation of phenomenological findings to a design of mixed-species stands and advanced silvicultural prescriptions by means of modelling. First, the knowledge of mixing effects on the structure and growth at the stand, species, and individual tree level is reviewed, with a focus on those findings that are most essential for suitable modelling and silvicultural designs and the regulation of mixed stands as opposed to monocultures. Then, the key role of growth models, stand simulators, and scenario assessments for designing mixed species stands is discussed. The next section illustrates that existing forest stand growth models require some fundamental modifications to become suitable for both monocultures and mixed-species stands. We then explore how silvicultural prescriptions derived from scenario runs would need to be both quantified and simplified for transfer to forest management and demonstrated in training plots. Finally, we address the main remaining knowledge gaps that could be remedied through empirical research.


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Toward managing mixed-species stands: from parametrization to prescription

Show Author's information Hans Pretzsch1( )Eric K. Zenner2
Chair for Forest Growth and Yield Science, Center of Life and Food Sciences Weihenstephan, Technical University of Munich, Hans-Carl-von-Carlowitz-Platz 2, 85354 Freising, Germany
Department of Forest Ecosystem Science and Management, Penn State University, University Park 16801, PA, USA

Abstract

A better understanding and a more quantitative design of mixed-species stands will contribute to more integrative and goal-oriented research in mixed-species forests. Much recent work has indicated that the structure and growth of mixed species forests may fundamentally differ from monocultures. Here we suggest how to progress from the present accumulation of phenomenological findings to a design of mixed-species stands and advanced silvicultural prescriptions by means of modelling. First, the knowledge of mixing effects on the structure and growth at the stand, species, and individual tree level is reviewed, with a focus on those findings that are most essential for suitable modelling and silvicultural designs and the regulation of mixed stands as opposed to monocultures. Then, the key role of growth models, stand simulators, and scenario assessments for designing mixed species stands is discussed. The next section illustrates that existing forest stand growth models require some fundamental modifications to become suitable for both monocultures and mixed-species stands. We then explore how silvicultural prescriptions derived from scenario runs would need to be both quantified and simplified for transfer to forest management and demonstrated in training plots. Finally, we address the main remaining knowledge gaps that could be remedied through empirical research.

Keywords: Multiplicative mixing effects, Overyielding, Overdensity, Modelling mixing effects, Scenario analysis, Silvicultural prescriptions, Practical guidelines

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Received: 01 July 2017
Accepted: 15 September 2017
Published: 10 October 2017
Issue date: December 2017

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© The Author(s) 2017.

Acknowledgements

Acknowledgements

The first author thanks the European Union for funding of the project "Management of mixed-species stands. Options for a low-risk forest management (REFORM)" (# 2816ERA02S), the Bavarian State Ministry for Nutrition, Agriculture, and Forestry for permanent support of the project W 07" Long-term experimental plots for forest growth and yield research "(# 7831-22209-2013), and the German Science Foundation for providing the funds for the projects PR 292/12-1" Tree and stand-level growth reactions on drought in mixed versus pure forests of Norway spruce and European beech". The second author thanks the National Institute of Food and Agriculture/Pennsylvania Agriculture Experiment Station project PEN 04516 for its support. Thanks are also to Ulrich Kern for the graphical artwork, Jeri Peck for initial review, and anonymous reviewers for improving the manuscript by their constructive criticism.

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